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headshot of John E. Titus

John E. Titus

Professor Emeritus

Biological Sciences


Titus' research interests lie in documenting compositional change in both ground-layer vegetation and trees, and experimentally testing the impacts of deer and soil acidity. He hopes these studies will help guide forest restoration projects.


  • PhD, MS, University of Wisconsin at Madison
  • BA, Oberlin College

Research Interests

  • Forest ecology and restoration